• 2007 - 2008: Postdoctoral Researcher (with Alice McHardy), Computational Genomics and Epidemiology, Max Planck Institut fuer Informatik, Saarbruecken, Germany.
  
  
• 2005 - 2007: Research Associate (JSPS, with Akira Sasaki), Department of Biology, Kyushu University, Fukuoka, Japan.
  
  
• 2003 - 2005: Research Associate (Wellcome Trust, with Mike Boots), Department of Animal & Plant Science, University of Sheffield, Sheffield, UK.
  
  
• 2000 - 2003: PhD (EPSRC, with Andy White and Tim Lenton), Department of Mathematics, Heriot-Watt University and Centre for Ecology and Hydrology, Edinburgh, UK.